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In this Letter, we study the collective behaviour of a large number of self-propelled microswimmers immersed in a fluid. Using unprecedently large-scale lattice Boltzmann simulations, we reproduce the transition to bacterial turbulence. We show that, even well below the transition, swimmers move in a correlated fashion that cannot be described by a mean-field approach. We develop a novel kinetic theory that captures these correlations and is non-perturbative in the swimmer density. To provide an experimentally accessible measure of correlations, we calculate the diffusivity of passive tracers and reveal its non-trivial density dependence. The theory is in quantitative agreement with the lattice Boltzmann simulations and captures the asymmetry between pusher and puller swimmers below the transition to turbulence.
Microswimmers exhibit an intriguing, highly-dynamic collective motion with large-scale swirling and streaming patterns, denoted as active turbulence -- reminiscent of classical high-Reynolds-number hydrodynamic turbulence. Various experimental, numer
Tracer particles immersed in suspensions of biological microswimmers such as E. coli or Chlamydomonas display phenomena unseen in conventional equilibrium systems, including strongly enhanced diffusivity relative to the Brownian value and non-Gaussia
We develop a statistical framework for the rheology of dense, non-Brownian suspensions, based on correlations in a space representing forces, which is dual to position space. Working with the ensemble of steady state configurations obtained from simu
Suspensions of rear- and front-actuated microswimmers immersed in a fluid, known respectively as ``pushers and ``pullers, display qualitatively different collective behaviours: beyond a characteristic density, pusher suspensions exhibit a hydrodynami
Dense suspensions of particles are relevant to many applications and are a key platform for developing a fundamental physics of out-of-equilibrium systems. They present challenging flow properties, apparently turning from liquid to solid upon small c