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We present theory and experiments demonstrating the existence of invariant manifolds that impede the motion of microswimmers in two-dimensional fluid flows. One-way barriers are apparent in a hyperbolic fluid flow that block the swimming of both smooth-swimming and run-and-tumble emph{Bacillus subtilis} bacteria. We identify key phase-space structures, called swimming invariant manifolds (SwIMs), that serve as separatrices between different regions of long-time swimmer behavior. When projected into $xy$-space, the edges of the SwIMs act as one-way barriers, consistent with the experiments.
We consider self-propelled droplets which are driven by internal flow. Tracer particles, which are advected by the flow, in general follow chaotic trajectories, even though the motion of the autonomous swimmer is completely regular. The flow is mixin
In an incompressible flow, fluid density remains invariant along fluid element trajectories. This implies that the spatial distribution of non-interacting noninertial particles in such flows cannot develop density inhomogeneities beyond those that ar
Transport and mixing of scalar quantities in fluid flows is ubiquitous in industry and Nature. Turbulent flows promote efficient transport and mixing by their inherent randomness. Laminar flows lack such a natural mixing mechanism and efficient trans
The transport of small quantities of liquid on a solid surface is inhibited by the resistance to motion caused by the contact between the liquid and the solid. To overcome such resistance, motion can be externally driven through gradients in electric
We investigate the role of linear mechanisms in the emergence of nonlinear horizontal self-propelled states of a heaving foil in a quiescent fluid. Two states are analyzed: a periodic state of unidirectional motion and a quasi-periodic state of slow