No Arabic abstract
We study the dynamics arising when two identical oscillators are coupled near a Hopf bifurcation where we assume a parameter $epsilon$ uncouples the system at $epsilon=0$. Using a normal form for $N=2$ identical systems undergoing Hopf bifurcation, we explore the dynamical properties. Matching the normal form coefficients to a coupled Wilson-Cowan oscillator network gives an understanding of different types of behaviour that arise in a model of perceptual bistability. Notably, we find bistability between in-phase and anti-phase solutions that demonstrates the feasibility for synchronisation to act as the mechanism by which periodic inputs can be segregated (rather than via strong inhibitory coupling, as in existing models). Using numerical continuation we confirm our theoretical analysis for small coupling strength and explore the bifurcation diagrams for large coupling strength, where the normal form approximation breaks down.
For many physical systems the transition from a stationary solution to sustained small amplitude oscillations corresponds to a Hopf bifurcation. For systems involving impacts, thresholds, switches, or other abrupt events, however, this transition can be achieved in fundamentally different ways. This paper reviews 20 such `Hopf-like bifurcations for two-dimensional ODE systems with state-dependent switching rules. The bifurcations include boundary equilibrium bifurcations, the collision or change of stability of equilibria or folds on switching manifolds, and limit cycle creation via hysteresis or time delay. In each case a stationary solution changes stability and possibly form, and emits one limit cycle. Each bifurcation is analysed quantitatively in a general setting: we identify quantities that govern the onset, criticality, and genericity of the bifurcation, and determine scaling laws for the period and amplitude of the resulting limit cycle. Complete derivations based on asymptotic expansions of Poincare maps are provided. Many of these are new, done previously only for piecewise-linear systems. The bifurcations are collated and compared so that dynamical observations can be matched to geometric mechanisms responsible for the creation of a limit cycle. The results are illustrated with impact oscillators, relay control, automated balancing control, predator-prey systems, ocean circulation, and the McKean and Wilson-Cowan neuron models.
In this work we have investigated the evolutionary dynamics of a generalist pathogen, e.g. a virus population, that evolves towards specialisation in an environment with multiple host types. We have particularly explored under which conditions generalist viral strains may rise in frequency and coexist with specialist strains or even dominate the population. By means of a nonlinear mathematical model and bifurcation analysis, we have determined the theoretical conditions for stability of nine identified equilibria and provided biological interpretation in terms of the infection rates for the viral specialist and generalist strains. By means of a stability diagram we identified stable fixed points and stable periodic orbits, as well as regions of bistability. For arbitrary biologically feasible initial population sizes, the probability of evolving towards stable solutions is obtained for each point of the analyzed parameter space. This probability map shows combinations of infection rates of the generalist and specialist strains that might lead to equal chances for each type becoming the dominant strategy. Furthermore, we have identified infection rates for which the model predicts the onset of chaotic dynamics. Several degenerate Bogdanov-Takens and zero-Hopf bifurcations are detected along with generalized Hopf and zero-Hopf bifurcations. This manuscript provides additional insights into the dynamical complexity of host-pathogen evolution towards different infection strategies.
This paper concerns two-dimensional Filippov systems --- ordinary differential equations that are discontinuous on one-dimensional switching manifolds. In the situation that a stable focus transitions to an unstable focus by colliding with a switching manifold as parameters are varied, a simple sufficient condition for a unique local limit cycle to be created is established. If this condition is violated, three nested limit cycles may be created simultaneously. The result is achieved by constructing a Poincare map and generalising analytical arguments that have been employed for continuous systems. Necessary and sufficient conditions for the existence of pseudo-equilibria (equilibria of sliding motion on the switching manifold) are also determined. For simplicity only piecewise-linear systems are considered.
In this paper, we study the existence and the property of the Hopf bifurcation in the two-strategy replicator dynamics with distributed delays. In evolutionary games, we assume that a strategy would take an uncertain time delay to have a consequence on the fitness (or utility) of the players. As the mean delay increases, a change in the stability of the equilibrium (Hopf bifurcation) may occur at which a periodic oscillation appears. We consider Dirac, uniform, Gamma, and discrete delay distributions, and we use the Poincare- Lindstedts perturbation method to analyze the Hopf bifurcation. Our theoretical results are corroborated with numerical simulations.
We describe combinatorial approaches to the question of whether families of real matrices admit pairs of nonreal eigenvalues passing through the imaginary axis. When the matrices arise as Jacobian matrices in the study of dynamical systems, these conditions provide necessary conditions for Hopf bifurcations to occur in parameterised families of such systems. The techniques depend on the spectral properties of additive compound matrices: in particular, we associate with a product of matrices a signed, labelled digraph termed a DSR^[2] graph, which encodes information about the second additive compound of this product. A condition on the cycle structure of this digraph is shown to rule out the possibility of nonreal eigenvalues with positive real part. The techniques developed are applied to systems of interacting elements termed interaction networks, of which networks of chemical reactions are a special case.