A system with charge conservation and lattice translation symmetry has a well-defined filling $ u$, which is a real number representing the average charge per unit cell. We show that if $ u$ is fractional (i.e. not an integer), this imposes very strong constraints on the low-energy theory of the system and give a framework to understand such constraints in great generality, vastly generalizing the Luttinger and Lieb-Schultz-Mattis theorems. The most powerful constraint comes about if $ u$ is continuously tunable (i.e. the system is charge-compressible), in which case we show that the low-energy theory must have a very large emergent symmetry group -- larger than any compact Lie group. An example is the Fermi surface of a Fermi liquid, where the charge at every point on the Fermi surface is conserved. We expect that in many, if not all, cases, even exotic non-Fermi liquids will have the same emergent symmetry group as a Fermi liquid, even though they could have very different dynamics. We call a system with this property an ersatz Fermi liquid. We show that ersatz Fermi liquids share a number of properties in common with Fermi liquids, including Luttingers theorem (which is thus extended to a large class of non-Fermi liquids) and periodic quantum oscillations in the response to an applied magnetic field. We also establis