(abridged) In this paper we derive observed and modelled shape parameters (apparent ellipticity and orientation of the ellipse) of 650 Galactic open clusters identified in the ASCC-2.5 catalogue. We provide the observed shape parameters of Galactic open clusters, computed with the help of a multi-component analysis. For the vast majority of clusters these parameters are determined for the first time. High resolution (star by star) N-body simulations are carried out with the specially developed $phi$GRAPE code providing models of clusters of different initial masses, Galactocentric distances and rotation velocities. The comparison of models and observations of about 150 clusters reveals ellipticities of observed clusters which are too low (0.2 vs. 0.3), and offers the basis to find the main reason for this discrepancy. The models predict that after $approx 50$ Myr clusters reach an oblate shape with an axes ratio of $1.65:1.35:1$, and with the major axis tilted by an angle of $q_{XY} approx 30^circ$ with respect to the Galactocentric radius due to differential rotation of the Galaxy. Unbiased estimates of cluster shape parameters require reliable membership determination in large cluster areas up to 2-3 tidal radii where the density of cluster stars is considerably lower than the background. Although dynamically bound stars outside the tidal radius contribute insignificantly to the cluster mass, their distribution is essential for a correct determination of cluster shape parameters. In contrast, a restricted mass range of cluster stars does not play such a dramatic role, though deep surveys allow to identify more cluster members and, therefore, to increase the accuracy of the observed shape parameters.
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