The blazar 3C454.3 exhibited a strong flare seen in gamma-rays, X-rays, and optical/NIR bands during 3--12 December 2009. Emission in the V and J bands rose more gradually than did the gamma-rays and soft X-rays, though all peaked at nearly the same time. Optical polarization measurements showed dramatic changes during the flare, with a strong anti-correlation between optical flux and degree of polarization (which rose from ~ 3% to ~ 20%) during the declining phase of the flare. The flare was accompanied by large rapid swings in polarization angle of ~ 170 degree. This combination of behaviors appear to be unique. We have cm-band radio data during the same period but they show no correlation with variations at higher frequencies. Such peculiar behavior may be explained using jet models incorporating fully relativistic effects with a dominant source region moving along a helical path or by a shock-in-jet model incorporating three-dimensional radiation transfer if there is a dominant helical magnetic field. We find that spectral energy distributions at different times during the flare can be fit using modified one-zone models where only the magnetic field strength and particle break frequencies and normalizations need change. An optical spectrum taken at nearly the same time provides an estimate for the central black hole mass of ~ 2.3 * 10^9 M_sun. We also consider two weaker flares seen during the $sim 200$ d span over which multi-band data are available. In one of them, the V and J bands appear to lead the $gamma$-ray and X-ray bands by a few days; in the other, all variations are simultaneous.
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