Experimentally observed ground state band based on the $1/2^{-}[521]$ Nilsson state and the first exited band based on the $7/2^{-}[514]$ Nilsson state in the odd-$Z$ nucleus $^{255}$Lr are studied by the cranked shell model (CSM) with the paring correlations treated by the particle-number-conserving (PNC) method. This is the first time the detailed theoretical investigations being performed on these rotational bands. Both the experimental kinematic and dynamic moment of inertia ($mathcal{J}^{(1)}$ and $mathcal{J}^{(2)}$) versus rotational frequency are reproduced quite well by the PNC-CSM calculations. By comparing the theoretical kinematic moment of inertia $mathcal{J}^{(1)}$ with the experimental ones extracted from different spin assignments, the spin $17/2^{-}rightarrow13/2^{-}$ is assigned to the lowest-lying $196.6(5)$ keV transition of the $1/2^{-}[521]$ band, and $15/2^{-}rightarrow11/2^{-}$ to the $189(1)$ keV transition of the $7/2^{-}[514]$ band, respectively. The proton $N=7$ major shell is included in the calculations. The intruder of the high$-j$ low$-Omega$ orbitals $1j_{15/2}$ $ (1/2^{-}[770])$ at the high spin leads to the band-crossing at $hbaromegaapprox0.20$ ($hbaromegaapprox0.25$) MeV for the $7/2^{-}[514]$ $alpha=-1/2$ ($alpha=+1/2$) band, and at $hbaromegaapprox0.175$ MeV for the $1/2^{-}[521]$ $alpha=-1/2$ band, respectively. Further investigations show that the band-crossing frequencies are quadrupole deformation dependent.
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