We present the analysis of the emission line galaxies members of 46 low redshift (0.04 < z < 0.07) clusters observed by WINGS (WIde-field Nearby Galaxy cluster Survey, Fasano et al. 2006). Emission line galaxies were identified following criteria that are meant to minimize biases against non-star forming galaxies and classified employing diagnostic diagrams. We have examined the emission line properties and frequencies of star forming galaxies, transition objects and active galactic nuclei (AGNs: LINERs and Seyferts), unclassified galaxies with emission lines, and quiescent galaxies with no detectable line emission. A deficit of emission line galaxies in the cluster environment is indicated by both a lower frequency with respect to control samples, and by a systematically lower Balmer emission line equivalent width and luminosity (up to one order of magnitude in equivalent width with respect to control samples for transition objects) that implies a lower amount of ionised gas per unit mass and a lower star formation rate if the source is classified as Hii region. A sizable population of transition objects and of low-luminosity LINERs (approx. 10 - 20% of all emission line galaxies) is detected among WINGS cluster galaxies. With respect to Hii sources they are a factor of approx. 1.5 more frequent than (or at least as frequent as) in control samples. Transition objects and LINERs in cluster are most affected in terms of line equivalent width by the environment and appear predominantly consistent with retired galaxies. Shock heating can be a possible gas excitation mechanism able to account for observed line ratios. Specific to the cluster environment, we suggest interaction between atomic and molecular gas and the intracluster medium as a possible physical cause of line-emitting shocks.