Sources at the brightest end of QSO luminosity function during the peak epoch of star formation and black hole accretion (z~2-4, i.e. Cosmic noon) are privileged sites to study the feeding & feedback cycle of massive galaxies. We perform the first systematic study of cold gas properties in the most luminous QSOs, by characterising their host-galaxies and environment. We analyse ALMA, NOEMA and JVLA observations of FIR continuum, CO and [CII] emission lines in eight QSOs ($L_{rm Bol}>3times10^{47}$ erg/s) from the WISSH sample at z~2.4-4.7. We report a 100% emission line detection rate and a 80% detection rate in continuum emission, and we find CO emission to be consistent with the steepest CO ladders observed so far. Sub-mm data reveal presence of (one or more) bright companion galaxies around 80% of WISSH QSOs, at projected distances of 6-130 kpc. We observe a variety of sizes for the molecular gas reservoirs (1.7-10 kpc), associated with rotating disks with disturbed kinematics. WISSH QSOs typically show lower CO luminosity and higher star formation efficiency than FIR matched, z~0-3 main-sequence galaxies, implying that, given the observed SFR ~170-1100 $M_odot$/yr, molecular gas is converted into stars on <50 Myr. Most targets show extreme dynamical to black-hole mass ratios $M_{rm dyn}/M_{rm BH}sim3-10$, two orders of magnitude smaller than local relations. The molecular gas fraction in WISSH hosts is lower by a factor of ~10-100 than in star forming galaxies with similar $M_*$. WISSH QSOs undergo an intense growth phase of both the central SMBH and host-galaxy. They pinpoint high-density sites where giant galaxies assemble and mergers play a major role in the build-up of the final host-galaxy mass. The observed low molecular gas fraction and short depletion timescale are likely due to AGN feedback, as traced by fast AGN-driven ionised outflows in all our targets.
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