We study the dynamics of nonlinear random walks on complex networks. We investigate the role and effect of directed network topologies on long-term dynamics. While a period-doubling bifurcation to alternating patterns occurs at a critical bias parameter value, we find that some directed structures give rise to a different kind of bifurcation that gives rise to quasi-periodic dynamics. This does not occur for all directed network structure, but only when the network structure is sufficiently directed. We find that the onset of quasi-periodic dynamics is the result of a Neimark-Sacker bifurcation, where a pair of complex-conjugate eigenvalues of the system Jacobian passes through the unit circle, destabilizing the stationary distribution with high-dimensional rotations. We investigate the nature of these bifurcations, study the onset of quasi-periodic dynamics as network structure is tuned to be more directed, and present an analytically tractable case of a four-neighbor ring.
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