The axion is a hypothetical, well-motivated dark-matter particle whose existence would explain the lack of charge-parity violation in the strong interaction. In addition to this original motivation, an `axiverse of ultra-light axions (ULAs) with masses $10^{-33},{rm eV}lesssim m_{rm a}lesssim 10^{-10},{rm eV}$ also emerges from string theory. Depending on the mass, such a ULA contributes to the dark-matter density, or alternatively, behaves like dark energy. At these masses, ULAs classical wave-like properties are astronomically manifested, potentially mitigating observational tensions within the $Lambda$CDM paradigm on local-group scales. ULAs also provide signatures on small scales such as suppression of structure, interference patterns and solitons to distinguish them from heavier dark matter candidates. Through their gravitational imprint, ULAs in the presently allowed parameter space furnish a host of observational tests to target in the next decade, altering standard predictions for microwave background anisotropies, galaxy clustering, Lyman-$alpha$ absorption by neutral hydrogen along quasar sightlines, pulsar timing, and the black-hole mass spectrum.
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