We present new HST ACS medium- and narrow-band images and long-slit, optical (4000 - 7200A) spectra obtained using the Isaac Newton Telescope (INT) on La Palma, of the merging system Mrk273. The HST observations sample the [OIII]4959,5007 emission from the galaxy and the nearby continuum. The images show that the morphologies of the extended continuum and the ionised gas emission from the galaxy are decoupled, extending almost perpendicular to each other. In particular, we detect for the first time a spectacular structure of ionised gas in the form of filaments extending ~23 kpc to the east of the nuclear region. The quiescent ionised gas kinematics at these locations suggests that these filaments are tidal debris left over from a secondary merger event that are illuminated by an AGN in the nuclear regions. The images also reveal a complex morphology in the nuclear region of the galaxy for both the continuum and the [OIII] emission. Kinematic disturbance, in the form of broad (FWHM > 500 km s-1) and/or strongly shifted (abs(DeltaV) >150 km s-1) emission line components, is found at almost all locations within a radius of ~4 kpc to the east and west of the northern nucleus. We fit the profiles of all the emission lines of different ionisation with a kinematic model using up to 3 Gaussian components. From these fits we derive diagnostic line ratios that are used to investigate the ionisation mechanisms at the different locations in the galaxy. We show that, in general, the line ratios are consistent with photoionization by an AGN as the main ionisation mechanism. Finally, the highest surface brightness [OIII] emission is found in a compact region that is coincident with the so-called SE nuclear component. The compactness, kinematics and emission line ratios of this component suggest that it is a separate nucleus with its own AGN.