If the cosmological inflationary scenario took place in the cosmic landscape in string theory, the inflaton, the scalar mode responsible for inflation, would have meandered in a complicated multi-dimensional potential. We show that this meandering property naturally leads to many e-folds of inflation, a necessary condition for a successful inflationary scenario. This behavior also leads to fluctuations in the primordial power spectrum of the cosmic microwave background radiation, which may be detected in a near future cosmic variance limited experiment like PLANCK.