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An example of Brunet-Derrida behavior for a branching-selection particle system on $Z$

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 Added by Jean Berard
 Publication date 2008
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and research's language is English
 Authors Jean Berard




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We consider a branching-selection particle system on $Z$ with $N geq 1$ particles. During a branching step, each particle is replaced by two new particles, whose positions are shifted from that of the original particle by independently performing two random walk steps according to the distribution $p delta_{1} + (1-p) delta_{0}$, from the location of the original particle. During the selection step that follows, only the N rightmost particles are kept among the 2N particles obtained at the branching step, to form a new population of $N$ particles. After a large number of iterated branching-selection steps, the displacement of the whole population of $N$ particles is ballistic, with deterministic asymptotic speed $v_{N}(p)$. As $N$ goes to infinity, $v_{N}(p)$ converges to a finite limit $v_{infty}(p)$. The main result is that, for every $0<p<1/2$, as $N$ goes to infinity, the order of magnitude of the difference $v_{infty}(p)- v_{N}(p)$ is $log(N)^{-2}$. This is called Brunet-Derrida behavior in reference to the 1997 paper by E. Brunet and B. Derrida Shift in the velocity of a front due to a cutoff (see the reference within the paper), where such a behavior is established for a similar branching-selection particle system, using both numerical simulations and heuristic arguments.



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251 - Jean Berard 2010
We consider a class of branching-selection particle systems on $R$ similar to the one considered by E. Brunet and B. Derrida in their 1997 paper Shift in the velocity of a front due to a cutoff. Based on numerical simulations and heuristic arguments, Brunet and Derrida showed that, as the population size $N$ of the particle system goes to infinity, the asymptotic velocity of the system converges to a limiting value at the unexpectedly slow rate $(log N)^{-2}$. In this paper, we give a rigorous mathematical proof of this fact, for the class of particle systems we consider. The proof makes use of ideas and results by R. Pemantle, and by N. Gantert, Y. Hu and Z. Shi, and relies on a comparison of the particle system with a family of $N$ independent branching random walks killed below a linear space-time barrier.
We introduce particle systems in one or more dimensions in which particles perform branching Brownian motion and the population size is kept constant equal to $N > 1$, through the following selection mechanism: at all times only the $N$ fittest particles survive, while all the other particles are removed. Fitness is measured with respect to some given score function $s:R^d to R$. For some choices of the function $s$, it is proved that the cloud of particles travels at positive speed in some possibly random direction. In the case where $s$ is linear, we show under some assumptions on the initial configuration that the shape of the cloud scales like $log N$ in the direction parallel to motion but at least $c(log N)^{3/2}$ in the orthogonal direction for some $c > 0$. We conjecture that the exponent 3/2 is sharp. This result is equivalent to the following result of independent interest: in one-dimensional systems, the genealogical time is greater than $c(log N)^3$, thereby contributing a step towards the original predictions of Brunet and Derrida. We discuss several open problems and also explain how our results can be viewed as a rigorous justification of Weismanns arguments for the role of recombination in population genetics.
We consider a branching-selection system in $mathbb {R}$ with $N$ particles which give birth independently at rate 1 and where after each birth the leftmost particle is erased, keeping the number of particles constant. We show that, as $Ntoinfty$, the empirical measure process associated to the system converges in distribution to a deterministic measure-valued process whose densities solve a free boundary integro-differential equation. We also show that this equation has a unique traveling wave solution traveling at speed $c$ or no such solution depending on whether $cgeq a$ or $c<a$, where $a$ is the asymptotic speed of the branching random walk obtained by ignoring the removal of the leftmost particles in our process. The traveling wave solutions correspond to solutions of Wiener-Hopf equations.
Motivated by the goal of understanding the evolution of populations undergoing selection, we consider branching Brownian motion in which particles independently move according to one-dimensional Brownian motion with drift, each particle may either split into two or die, and the difference between the birth and death rates is a linear function of the position of the particle. We show that, under certain assumptions, after a sufficiently long time, the empirical distribution of the positions of the particles is approximately Gaussian. This provides mathematically rigorous justification for results in the biology literature indicating that the distribution of the fitness levels of individuals in a population over time evolves like a Gaussian traveling wave.
We establish rigorous upper and lower bounds for the speed of pulled fronts with a cutoff. We show that the Brunet-Derrida formula corresponds to the leading order expansion in the cut-off parameter of both the upper and lower bounds. For sufficiently large cut-off parameter the Brunet-Derrida formula lies outside the allowed band determined from the bounds. If nonlinearities are neglected the upper and lower bounds coincide and are the exact linear speed for all values of the cut-off parameter.
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