We investigate the impact of neutral hydrogen (HI) in galaxies on the statistics of 21-cm fluctuations using analytic and semi-numerical modelling. Following the reionisation of hydrogen the HI content of the Universe is dominated by damped absorption systems (DLAs), with a cosmic density in HI that is observed to be constant at a level equal to ~2% of the cosmic baryon density from z~1 to z~5. We show that extrapolation of this constant fraction into the reionisation epoch results in a reduction of 10-20% in the amplitude of 21-cm fluctuations over a range of spatial scales. The assumption of a different percentage during the reionisation era results in a proportional change in the 21-cm fluctuation amplitude. We find that consideration of HI in galaxies/DLAs reduces the prominence of the HII region induced shoulder in the 21-cm power spectrum (PS), and hence modifies the scale dependence of 21-cm fluctuations. We also estimate the 21cm-galaxy cross PS, and show that the cross PS changes sign on scales corresponding to the HII regions. From consideration of the sensitivity for forthcoming low-frequency arrays we find that the effects of HI in galaxies/DLAs on the statistics of 21-cm fluctuations will be significant with respect to the precision of a PS or cross PS measurement. In addition, since overdense regions are reionised first we demonstrate that the cross-correlation between galaxies and 21-cm emission changes sign at the end of the reionisation era, providing an alternative avenue to pinpoint the end of reionisation. The sum of our analysis indicates that the HI content of the galaxies that reionise the universe will need to be considered in detailed modelling of the 21-cm intensity PS in order to correctly interpret measurements from forthcoming low-frequency arrays.
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